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The main source of primary infection comes from hypha, akinetes and sclerotiums over-wintering in the bulb, residues of diseased plants and the soil. 病菌以菌丝体在种球内，或以菌丝体、厚垣袍子及菌核随病残体在土壤中越冬，成为翌年的主要初侵染源。