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The Cirststep is catalyzed by choi inc monooxygenase(CMO), the second byhetai ne aidehyde dehydrogonase(BADII). 其中催化第一步反應的酶爲膽堿單氧化物酶（CMO），第二步反應由甜菜堿醛脫氫酶（BADH）催化。