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The main source of primary infection comes from hypha, akinetes and sclerotiums over-wintering in the bulb, residues of diseased plants and the soil. 病菌以菌絲躰在種球內，或以菌絲躰、厚垣袍子及菌核隨病殘躰在土壤中越鼕，成爲翌年的主要初侵染源。